A new play mechanic called 'Band Moments' will require all members of the band to play sections of a song successfully to gain rewards, both in scoring and visual effects on screen. Star Power is no longer shared by the band, and instead tracked for each player in a similar manner as Rock Band. While playing in a band, Star Power is now tracked separately for each player, as opposed to collectively for the band as in World Tour.

Whereas in World Tour, a band could only have one of each instrument, Guitar Hero 5 allows players to arrange for any combination of instruments, including all four players on the same instrument if they so choose. Players can also play in groups of up to four local or remote players to form a band, co-operatively playing through a song. Specially marked sections of the song, if completed correctly, help to build up Star Power when then can be unleashed through an action with the controller to further double the current score multiplier.ДЄs with Guitar Hero World Tour, Guitar Hero 5 supports the playing of lead and bass guitar through guitar controllers, drums through a drum controller, and vocals through a microphone.

Correctly hitting consecutive notes will build the player's score multiplier. Should the meter fall below a certain threshold, the song will end prematurely with the player booed off the stage by a virtual audience.

Hitting correct notes in time with the music increases the player's score and builds their performance meter, while missing notes will cause the meter to drop. Using a special game controller, players attempt to match scrolling notes as they appear on screen to mimic the playing of rock music and other songs. Gameplay in Guitar Hero 5 is similar to previous games in the series.

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